1991-1995
Undergraduate: Oakland University, B.S. Biochemistry, minor in Chemistry
Mentor: Dr. Barry Winkler

1995-2001
Graduate:University of California Santa Barbara, Ph.D. Molecular, Cellular and Developmental Biology
Mentor: Dr. Steven Fisher

2001-2007
Postgraduate Training: Postdoctoral fellowship in Retinal Gene Therapy, University of Pennsylvania
Mentor: Dr. Jean Bennett